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The yak ( Bos grunniens), also known as the Tartary ox, grunting ox, hairy cattle, sarlak or sarlyk, or domestic yak, is a species of long-haired found throughout the region, the , , the , and as far north as and . It is descended from the ( Bos mutus).


Etymology
The English word yak originates from the . In Tibetan and it refers only to the male of the species, the female being called or in Tibetan and in Balti. In English, as in most other languages that have borrowed the word, yak is usually used for both sexes, with bull or cow referring to each sex separately.


Taxonomy
Belonging to the genus , yaks are related to and . Mitochondrial DNA analyses to determine the evolutionary history of yaks have been inconclusive.

The yak may have diverged from cattle at any point between one and five million years ago, and there is some suggestion that it may be more closely related to bison than to the other members of its designated genus. Apparent close fossil relatives of the yak, such as , have been found in eastern Russia, suggesting a possible route by which yak-like ancestors of the modern could have entered the Americas.

The species was originally designated as Bos grunniens ("grunting ox") by in 1766. Still, this name is now generally considered to refer only to the domesticated form of the animal, with Bos mutus ("mute ox") being the preferred name for the wild species. Although some authors still consider the wild yak to be a , Bos grunniens mutus, the ICZN made an official ruling in 2003 permitting the use of the name Bos mutus for , and this is now the more common usage.

There are no recognised subspecies of yak except where the wild yak is considered a subspecies of Bos grunniens.


Physical characteristics
Yaks are heavily built animals with bulky frames, sturdy legs, rounded, cloven hooves, and extremely dense, long fur hanging lower than the belly. While wild yaks are generally dark, blackish to brown in colouration, domestic yaks can be quite variable, often having rusty brown and cream patches. They have small ears and broad foreheads, with smooth horns that are generally dark in colour. In males (bulls), the horns sweep out from the sides of the head and then curve backwards; they typically range from in length.

The horns of females (cows) are smaller, at in length, and have a more upright shape. Both sexes have a short neck with a pronounced hump over the shoulders, although this is larger and more visible in males. Males weigh , females weigh . Wild (feral) yaks can be substantially heavier, bulls reaching weights of up to .Buchholtz, C. (1990). True Cattle (Genus Bos). pp. 386–397 in S. Parker, ed. Grzimek's Encyclopedia of Mammals, Volume 5. New York: McGraw-Hill Publishing Company. (quoted in Oliphant, M. (2003). Bos grunniens (On-line), Animal Diversity Web. Accessed 4 April 2009) Depending on the breed, domestic yak males are high at the withers, while females are high at the withers.

Both sexes have long, shaggy hair with a dense woolly undercoat over the chest, flanks, and thighs to insulate them from the cold. Especially in bulls, this may form a long "skirt" that can reach the ground. The tail is long and horselike rather than tufted like the tails of cattle or bison. Domesticated yaks have a wide range of coat colours, with some individuals being white, grey, brown, roan or . The in females and the in males are small and hairy as protection against the cold. Females have four .

Yaks are not known to produce the characteristic lowing (mooing) sound of cattle, but both wild and domestic yaks grunt and squeak, which inspired the scientific name of the domestic yak variant, Bos grunniens (grunting bull). Nikolay Przhevalsky named the wild variant Bos mutus (silent bull), believing that it did not make a sound at all, but it does.


Physiology
Yak physiology is well adapted to high altitudes, having larger lungs and heart than cattle found at lower altitudes, as well as greater capacity for transporting oxygen through their blood, The Yak, Second Edition. Bangkok: Regional Office for Asia and the Pacific Food and Agriculture Organization of the United Nations, . Accessed 8 August 2008. due to the persistence of foetal haemoglobin throughout life. Conversely, yaks have trouble thriving at lower altitudes, Yak , Animal genetics training resources version II: Breed Information. Adopted from: Bonnemaire, J. "Yak". In: Mason, Ian L. (ed). (1984). Evolution of Domesticated Animals. London: Longman, pp. 39–45. . Accessed 8 August 2008. and are prone to suffering from heat exhaustion above about . Further adaptations to the cold include a thick layer of and an almost complete lack of functional .

Compared with domestic cattle, the of yaks is unusually large, relative to the . This likely allows them to consume greater quantities of low-quality food at a time, and to ferment it longer to extract more nutrients. Yak consume the equivalent of 1% of their body weight daily while cattle require 3% to maintain condition. They are grazing herbivores, with their wild ancestors feeding primarily on and , with some herbs and dwarf shrubs.


Reproduction and life history
Yaks mate in the summer, typically between July and September, depending on the local environment. For the remainder of the year, many bulls wander in small bachelor groups away from the large herds. Still, as the rut approaches, they become aggressive and regularly fight with each other to establish dominance. In addition to non-violent threat displays, bellowing, and scraping the ground with their horns, bull yaks compete more directly, repeatedly charging at each other with heads lowered or sparring with their horns. Like , but unlike cattle, males wallow in dry soil during the rut, often while scent-marking with urine or dung. Females enter oestrus up to four times a year, and females are receptive only for a few hours in each cycle.

lasts between 257 and 270 days, so that the young are born between May and June, and results in the birth of a single calf. The cow finds a secluded spot to give birth, but the calf can walk within about ten minutes of birth, and the pair soon rejoin the herd. Females of both the wild and domestic forms typically give birth only once every other year, although more frequent births are possible if the food supply is good.

Calves are at one year and become independent shortly thereafter. Wild calves are initially brown in color and only later develop darker adult hair. Females generally give birth for the first time at three or four years of age, and reach their peak reproductive fitness at around six years. Yaks may live for more than twenty years in domestication or captivity, although it is likely that this may be somewhat shorter in the wild.


Husbandry
For thousands of years, domesticated yaks have been kept in Mongolia and Tibet, primarily for their milk, (wool), and meat, and as . Their dried droppings are an important fuel, used all over Tibet, and are often the only fuel available on the high, treeless . Yaks transport goods across mountain passes for local farmers and traders and are an attraction for climbing and trekking expeditions: "Only one thing makes it hard to use yaks for long journeys in barren regions. They will not eat , which could be carried on the journey. They will starve unless they can be brought to a place where there is grass."Golden Book Encyclopedia, Vol. 16 p. 1505b. Rockefeller Center, NY: (1959). They also are used to draw . Yaks' milk is often processed to a cheese called in Tibetan and Nepali languages, and byaslag in Mongolia. is an ingredient of the that Tibetans consume in large quantities, and is also used in lamps and made into used in religious festivities." Butter - Yaks, butter & lamps in Tibet", WebExhibits.


Outside the Himalayas
Small numbers of herds can be found in the United States, Canada, New Zealand, and some parts of Europe. Yaks have generated interest outside the Himalayas as a commercial crop and by cattle breeders. The main interest of North American yak breeders is lean meat production by hybridizing with other cattle, followed by production.


Research
The Indian government established a dedicated centre for research into yak husbandry, the ICAR-National Research Centre on Yak, in 1989. It is located at , Arunachal Pradesh, and maintains a yak farm in the Nyukmadung area at an altitude of above MSL.


Breeding and hybridization
In , , and , domestic cattle are crossbred with yaks. This gives rise to the infertile male མཛོ། as well as fertile females known as dzomo or zhom མཛོ་མོ།, which may be crossed again with cattle. The Dwarf Lulu breed, "the only Bos primigenius taurus type of cattle in Nepal" has been tested for DNA markers and found to be a mixture of both taurine and types of cattle ( B. p. taurus and B. p. indicus) with yak. According to the International Veterinary Information Service, the low productivity of second-generation cattle–yak crosses makes them suitable only as meat animals.

Crosses between yaks and domestic cattle ( Bos primigenius taurus) have been recorded in Chinese literature for at least 2,000 years. Successful crosses have also been recorded between ,Zhang, R.C. (14 December 2000). "Interspecies Hybridization between Yak, Bos taurus and Bos indicus and Reproduction of the Hybrids". In: Recent Advances in Yak Reproduction, Zhao, X.X.; Zhang, R.C. (eds.). International Veterinary Information Service. , and , generally with similar results to those produced with domestic cattle.


Domestication
Jacques et al. (2021) show that most elaborate yak-related terminologies are found within Tibetic and languages. Both branches also have native terms for , suggesting that Tibetic and Gyalrongic speakers may have independently cross-bred yaks and cattle, predating the proto-Gyalrongic split (3221 2169–4319 ) from Tibeto-Gyalrongic. The oldest dated physical evidence of yak domestication is from 2,500 years BP.


Customs
In parts of Tibet and Karakorum, is a form of entertainment at traditional festivals and an important part of their culture. More recently, sports involving domesticated yaks, such as or , are being marketed as tourist attractions in South Asian countries, including in , Pakistan.

In Nepal, an annual festival is held to drink the fresh blood of yak, and it is believed that it cures various diseases such as gastritis, , and body strain. The fresh blood is extracted from the neck of a yak without killing it. The cut is healed after the ceremony is over. The ritual is believed to be originated in Tibet and .

Traditionally, yaks are used to transport deceased people to for in Tibet, with Tibetan practice holding that the yaks be set free upon the conclusion of the task.


Gallery
File:Yaks in Manali.jpg|Yaks in Manali, Himachal Pradesh, India, saddled for riding File:LitangMonastery.jpg|Train of pack yaks at Litang monastery in Garzê Tibetan Autonomous Prefecture, Sichuan, China File:Yaks still provide the best way to plow fields in Tibet.jpg|Yaks plowing fields in Tibet File:YakRace.jpg|Yaks in , Pakistan File:四川 阿坝 白牦牛 - panoramio.jpg|Domestic yak in , China File:四姑娘山景区 Mount Siguniang Scenic Area 17.jpg|A yak at , Sichuan, China File:Yak rearing at highest altitude- Rigma, Dolpa.jpg|Yak are reared on grazing pastures and meadows at elevations of 3,000–5,000 masl in 28 northern mountain districts in Nepal.


See also


External links

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